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[[Image:Wallace03.jpg|thumb|300px|A map showing the zoogeographical regions.<ref name=geog/>]]

'''Biogeography''' is the study of how [[species]] are distributed. It notes where organisms live, and why they are (or are not) found in a certain geographical area.

Biogeography teaches how animals and plants are [[adaptation|adapted]] to the places they live in, and how similar places often have quite different animals and plants.

Between about 1800 to 1855, [[natural history|natural historians]] made lists of species in various regions of the world. These lists were published as tables in their books.<ref name=von/><ref name=Al/> [[Charles Darwin]] and [[Alfred Russel Wallace]] published the idea of [[evolution]] by [[natural selection]]. They travelled to [[tropical]] countries, and wrote about the life in those countries.<ref name=geog/><ref name=Origin>Darwin, Charles 1859. ''[[On the Origin of Species]] by means of natural selection, or the preservation of favoured races in the struggle for life''. Murray, London. Chapters 10 and 11</ref> They said that evolution was the key to understanding geographical distribution.

New species are usually formed by [[speciation]] {{ndash}} an earlier species splitting into two. These species may travel to new places. but they may be stopped from travelling by mountains and seas, and by climate. This means that ''two places with similar climate often have different kinds of animals and plants''. For example [[marsupial]]s, which live in Australia, are very different from the fauna in South America. The species on islands ([[Hawaii]],<ref>Carlquist, Sherwin 1965. ''Island life: a natural history of the islands of the world''. American Museum of Natural History.</ref><ref>Carlquist, Sherwin 1970. ''Hawaii: a natural history''. American Museum of Natural History.</ref> [[Galapagos]]) may be very different to species on mainland continents.

:"The more distant countries are, the more dissimilar are their animals and plants; and the nearer they are, the more similar are their animals and plants".<ref name=Sclater/> This is true, but only up to a point. There are many examples of species with huge ranges.

== History ==
[[File:Plos wilson.jpg|thumb|175px|Edward O. Wilson, biologist and conservationist]]

The scientific theory of biogeography grows out of the work of [[Alexander von Humboldt]] (1769–1859),<ref name=von>von Humboldt 1805. ''Essai sur la geographie des plantes; accompagne d'un tableau physique des régions equinoxiales''. Levrault, Paris.</ref> [[Hewett Cottrell Watson]] (1804–1881),<ref>Watson H.C. 1847–1859. ''Cybele Britannica: or British plants and their geographical relations''. Longman, London.</ref> [[Alphonse de Candolle]] (1806–1893),<ref name=Al>de Candolle, Alphonse 1855. ''Géographie botanique raisonnée'' &c. Masson, Paris.</ref> [[Alfred Russel Wallace]] (1823–1913),<ref name=geog>Wallace A.R. 1876. ''The geographical distribution of animals''. 2 vols, Macmillan, London.</ref> [[Philip Lutley Sclater]] (1829–1913),<ref name=Sclater>Sclater P.L. 1879. Geographical distribution of mammals. ''The Scientific Man: a weekly illustrated journal of science''. volume 1 #22, New York, August 23, 1879.</ref> and other biologists and explorers.<ref>Browne, Janet 1983. ''The secular ark: studies in the history of biogeography''. Yale University Press, New Haven. {{ISBN|0-300-02460-6}}</ref>

Wallace studied the distribution of flora and fauna in the [[Amazon Basin]] and the [[Malay Archipelago]] in the mid-19th century.<ref>Wallace A.R. 1853. ''Travels on the Amazon and the Rio Negro''. London.</ref><ref>Wallace A.R. ''The Malay Archipelago''. London.</ref> Wallace and Sclater saw biogeography as a source of support for the theory of evolution. Key findings, such as the sharp difference in fauna either side of the [[Wallace Line]],<ref name="Mayr44">{{Citation | last = Mayr | first = E. | title = Wallace's Line in the light of recent zoogeographic studies | journal = The Quarterly Review of Biology | volume = 19 | issue = 1 | pages = 1–14 | year = 1944 | url = https://www.jstor.org/stable/2808563}}</ref> can only be understood in this light. Otherwise, the field of biogeography would be a purely descriptive one.

Both Darwin and Wallace gave a great deal of attention to [[wikt:ocean|oceanic]] islands as offering examples of evolution, especially [[speciation]]. Darwin visited the [[Galapagos Islands]] and studied its [[fauna]]. Wallace spent years on the islands of S.E. Asia.<ref>Wallace A.R. 1880. ''Island life''. Macmillan, London.</ref> This interest was revived by ''The theory of island biogeography'' by [[Robert MacArthur]] and [[Edward Osborne Wilson|E.O. Wilson]] in 1967. They showed that the variation in species in a single area could be predicted if one knew the habitat area, immigration rate, and extinction rate.

It was realised that [[habitat fragmentation|habitat fragments]] are like islands. They can be investigated by the same methods. This spurred the development of [[conservation biology]].

[[Genome analysis]] allows scientists to test theories about the origin and dispersal of populations, such as island species. It allows biologists to test theories about where the species come from.

== Books ==
*Cox C.B. and Moore P.D. 2010. ''Biogeography: an ecological and evolutionary approach''. 8th ed, Wiley. {{ISBN|0-470-63794-3}} (standard text)

== See also ==
*[[Speciation#Geographical isolation]]
*[[Great American Interchange]]
*[[Ecology]]

== References ==
{{Reflist}}

[[Category:Branches of biology]]
[[Category:Branches of geography]]

Biogeography - Revision history

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